Ecological succession is the gradual
    process by which ecosystems change and develop over time.  
    For example, a bare patch of ground will not stay bare.
    It will rapidly be colonized by a variety of plants. 
    
      
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        A recently cleared patch of ground (in Britain).  | 
        The
        same ground 2 years later,  
        now covered in grasses and low flowering plants.  | 
       
     
    In the process of succession, the species present in an
    area will gradually change. 
    Succession takes place because the environmental
    conditions in a particular place change over time. Each species is adapted to thrive and
    compete best against other species under a very specific set of environmental conditions.
    If these conditions change, then the existing species will be replaced by a new set of
    species which are better adapted to the new conditions. 
    As an example, the environmental conditions present on
    the bare patch of ground above would have been quite different 2 years later. Some of
    these differences are highlighted below. 
    
      
        | Bare Ground | 
        Two
        Years Later | 
       
      
        No plant competition for light, space,
        nutrients or water. 
        Soil mobile and liable to erosion and
        loss. 
        A more extreme surface microclimate
        because the bare soil both absorbs and reflects heat more than soil covered in vegetation. 
         
        A drier environment because there is no plant cover to hold moisture above ground and
        little humus to hold it in the soil. 
        Lower nutrient levels in the soil.  | 
        Intense plant competition for space and
        other resources. 
        Soil bound by roots and plant cover. 
        The plant cover provides a certain
        amount of ground insulation from extremes of temperature. There are now also a variety of
        microclimates within the vegetation.  
        Plant cover and increasing humus levels
        help to retain water.  
         
        The nutrient levels in the soil will have increased.
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    The bare ground conditions favour pioneer
    plant species. These are often species which grow best where there is little competition
    for space and resources.  
    
      
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        Mosses are often pioneer
        species. Most moss species are low growing, carpeting the ground and with little height.
        As a result of this growth form, many mosses are unable to successfully compete for space
        amongst taller, dense ground cover.  This makes bare ground
        ideal for the establishment of a number of different moss species. These mosses then
        provide a microhabitat equivalent to a miniature forest for a variety of invertebrates
        such as mites and spiders. The moss also acts like a sponge when wet, in some cases
        providing a semi-aquatic microhabitat.   | 
       
     
    
      
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        Pioneer
        species are often also 'opportunist' species which are able to rapidly exploit a sudden
        new opening in ground plant cover.   
        Left: Dandelion 
          
        Typical examples in the UK would be dandelion, Foxglove and
        willowherb.  
          
          
        Seeds arrive, germinate and grow quickly, rapidly reproducing
        themselves before other slower-colonizing species arrive to outcompete them. 
        Along with the plants will come the animals which feed on them or
        use them for shelter. 
          
          
        Left: Bee on Foxglove  | 
       
      
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        Pioneer species are often characterized by having light
        seeds, which are easily transported by the wind. Many of them belong to the flowering
        plant family 'Compositae' Such plants produce large numbers of
        seeds with parachutes of fine hairs which help to keep them afloat in air currents. 
          
        (Left: Ragwort seed)  | 
       
     
      
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