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The Woodland Education Centre
The Heathland Restoration Project
Trialling different methods of management for heathland restoration.
Contents

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APPENDIX A. SPECIES LIST 1996 - 2001
including abundance, expressed as percentage frequency.

Summary

  • Percentage frequency = the percentage of the total number of quadrats sampled over the whole site which contain the species. For example, a percentage frequency of 100%, means that the species was found in all quadrats sampled.
  • Percentage frequencies have been rounded to the nearest whole number, except for those which are less than 1%.
  • A + sign indicates that the species was seen on the site, but was not found in any of the quadrats sampled.
  • A simple definition of a heathland is 'a community of dwarf shrubs where heathers predominate'. However, beyond this very broad description, heathland habitats come in a variety of subtly different forms, each with slightly different assemblages of plant species. The plant species present on any one heath will depend on geographical area, elevation, microclimate and soil characteristics (in particular the amount of soil water typically present). A few plants are characteristic of almost all heaths (e.g. Heather/Ling and gorses), while others may be characteristic of certain types of heath.

    Species often associated with heath habitats in one form or another are picked out in green text.

 

SPECIES

COMMON NAME

% Freq

MOSSES

1996

1997

1998

1999

2000 2001
Atrichum undulatum (Hedw.)

0.3

13 3 3
Brachythecium rutabulum (Hedw.) B.,S.& G.

+

+

Bryum microerythrocarpum C. Mull & Kindb.

+

Campylopus atrovirens De Not. 0.4
Campylopus introflexus (Hedw.) Brid.

0.8

+

1
* Campylopus paradoxus Wils.

1.6

5

2 0.8 0.7
* Campylopus pyriformis (Schultz) Brid.

+

Dicranella heteromalla (Hedw.) Schp.

+

1
* Dicranum scoparium Hedw.

+

Eurhynchium praelongum (Hedw.) B.& S.

2

7

52 75 64
Hypnum andoi (formerly H. mammillatum)

1

+

0.6
Hypnum cupressiforme  Hedw. 3 2
* Hypnum jutlandicum Holmen & Warncke

+

10

14 20 19
Isothecium myurum  Brid. 0.8
Mnium hornum Hedw.

+

Plagiothecium denticulatum  (Hedw.) B., S. & G. 4
Polytrichum formosum Hedw.

0.7

23

34

38 36 33
Rhytidiadelphus squarrosus (Hedw.) Warnst.

+

2
Sphagnum sp. 0.3
Thuidium tamarascinum (Hedw.) B.,S.& G.

17

20

42 45 41
LIVERWORTS
Lophocolea bidentata (L.) Dum. var. bidentata

+

FERNS
* Blechnum spicant (L.) Roth.

Hard Fern

3

4

5

6 5 6
Dryopteris carthusiana (Villars) H.P.Fuchs;

Narrow Buckler Fern

9 3 9
Dryopteris felix-mas  (L.)Schott.

Male Fern

0.8 0.4
Polypodium vulgare  L.

Common Polypody

0.3
* Pteridium aquilinum (L.) Kuhn.

Bracken

3

11

9

8 7 7
ANGIOSPERMS
LILIACEAE
Hyacinthoides non-scripta (L) Chouard ex Rothm.

Bluebell

27

10

18

15 19 20
ARACEAE
Arum maculatum L.

Lords & Ladies

+

0.4
GRAMINAE
* Agrostis canina L.

Velvet Bent

+

4

8

25 26 17
* Agrostis capillaris  L.

Common Bent

+

67

81

80 74 75
* Agrostis curtisii  Kerguelen

Bristle Bent

+

9

14

7 0.3 0.7
Agrostis stolonifera L.

Creeping Bent

+

21

16

3 2
Agrostis vinealis Schreb.

Brown Bent

1 0.7
Anisantha sterilis  (L.) Nev.

Barren Brome

0.3
* Anthoxanthum odoratum L.

Sweet Vernal Grass

2

0.4

4 0.4 2
Arrhenatherum elatius (L.)

False Oat Grass

4

5

1

2 3 6
Bromus ramosus Huds.

Hairy Brome

+

0.8
Deschampsia cespitosa (L.) Beauv.

Tufted Hair-grass

0.4

Dactylis glomerata L.

Cocksfoot

0.8
Holcus lanatus L.

Yorkshire Fog

51

59

80

73 67 58
Molinia caerulea  (L.) Moench

Purple Moor-grass

+
CYPERACEAE
* Carex binervis Sm.

Green-ribbed Sedge

10

22

17 13 19
Carex flacca Schreb.

Glaucous Sedge

4 4 4
Carex flava agg.

Yellow Sedges

4

5

4 2 8
Carex laevigata Sm.

Smooth-stalked Sedge

14

9

12

21 16 10
Carex pallescens L.

Pale Sedge

+

* Carex pilulifera L.

Pill Sedge

+

2

17

21 7 12
Carex sylvatica Huds.

Wood Sedge

+

3

10

10 15 9
Carex strigosa Huds.

Thin-spiked Wood Sedge

+

JUNCACEAE
Juncus acutiflorus Ehrh

Sharp-flowered Rush

+
Juncus conglomeratus L.

Compact Rush

33

28

21

27 26 18
* Juncus effusus L.

Soft Rush

13

6

10

5 4 4
* Luzula campestris (L.) DC.

Field Woodrush

+

5

8 17 13
* Luzula multiflora (Retz.) Lej.

Heath Woodrush

17

9

10

19 15 5
Luzula pilosa (L.) Willd.

Hairy Woodrush

+

+

8

1 2 6
RANUNCULACEAE
Anemone nemorosa L.

Wood Anemone

+

Ranunculus repens L.

Creeping Buttercup

8 2

0.7

2 5 6
HYPERICACEAE
Hypericum androsaemum L.

Tutsan

0.7

+

0.4

0.4 +
Hypericum perforatum L.

Perforate St John's Wort

+
* Hypericum pulchrum L.

Slender St John’s Wort

56

52

42

21 18 22
CARYOPHYLLACEAE
Silene dioica (L.) Clairv.

Red Campion

+

0.4
VIOLACEAE
Viola riviniana Rchb.

Common Dog-violet

31

33

46

46 53 42
OXALIDACEAE
Oxalis acetosella L.

Wood Sorrel

0.7

2

2

2 3 4
PAPILIONACEAE
Lotus pedunculatus Cav.

Greater Bird’s Foot Trefoil

8

10

15

16 19 18
Trifolium pratense L.

Red Clover

0.4

+

0.3
Trifolium repens L.

White (Dutch) Clover

+

* Ulex europaeus L.

Common Gorse

53

33

32

36 34 38
* Ulex gallii Planch.

Western Gorse

+

0.6 0.4
ROSACEAE
Crataegus monogyna Jacq.

Hawthorn

0.4
Fragaria vesca L.

Wild Strawberry

3 0.4 0.8
* Potentilla erecta (L.) Rausch

Tormentil

+

1

5

5 4 5
Potentilla sterilis (L.) Garcke

Barren Strawberry

1

2 0.9
Prunus sp.

Cherry

15

13

17

15 10 11
* Rubus fruticosus agg.

Bramble

36

46

54

56 56 67
Rubus idaeus L.

Raspberry

2

+

+

10 15
Sorbus aucuparia L.

Rowan

+

1

2

8 10 6
ONAGRACEAE
Chamaenerion angustifolium (L.) Scop.

Rosebay Willowherb

0.8

+

1

+
Circaea lutetiana L.

Enchanter’s Nightshade

0.7

+

2

2 1 5
Epilobium montanum L.

Broad-leaved Willowherb

0.7

+

0.4

0.8
URTICACEAE
Urtica dioica L.

Common Nettle

0.3
AQUIFOLIACEAE
Ilex aquifolium L.

Holly

3

2

2

2 0.4 0.9
ARALIACEAE
Hedera helix L.

Ivy

1

3

4

5 4 7
FAGACEAE
Castanea sativa Mill.

Sweet Chestnut

+
Fagus sylvatica L.

Beech

+

+

+

0.3
Quercus sp.

Oak

1

2

9 6 3
BETULACEAE
* Betula pendula Roth.

Silver Birch

21

36

37

33 30 29
CORYLACEAE
Corylus avellana L.

Hazel

1 0.8 0.3
SALICACEAE
Salix auritaL.

Eared Willow

1

4

3

2 3 1
Salix caprea agg.

Goat Willow/Great Sallow

0.4

0.7

1 2 1
ACERACEAE
Acer pseudoplanatus L.

Sycamore

11

1

0.6 0.4 +
ERICACEAE
* Calluna vulgaris (L.) Hull.

Heather (Ling)

6

11

12

22 17 22
* Erica cinerea L.

Bell Heather

0.8

2

3

7 5 6
Rhododendron ponticum L.

Rhododendron

0.3

1

2 1
PRIMULACEAE
Lysimachia nemorum L.

Yellow Pimpernel

19

14

17

20 22 23
Primula vulgaris Huds.

Primrose

1

0.4

2

2 2
OLEACEAE
Fraxinus excelsior L.

Ash

1

+
GENTIANACEAE
Centaurium erythraea Rafn

Common Centaury

2

0.4

+
SCROPHULARIACEAE
Digitalis purpurea L.

Foxglove

28

19

8

7 3 1
Scrophularia nodosa L.

Common Figwort

0.5

2 5 0.9
Veronica chamaedrys L.

Germander Speedwell

3

3

2

3 7 5
Veronica montana L.

Wood Speedwell

4

0.3
Veronica officinalis L.

Heath Speedwell

10

12

16

14 17 19
LABIATEAE
Ajuga reptans L.

Bugle

6

10

10

19 23 24
Prunella vulgaris L.

Selfheal

+

0.4
Stachys sylvatica L.

Hedge Woundwort

3

3

9

9 12 9
Teucrium scorodonia L.

Wood Sage

6

10

11

9 8 12
CAPRIFOLIACEAE
Leycesteria formosa Wallich

Himalayan Honeysuckle

0.4
Lonicera periclymenum L.

Honeysuckle

2

1

1 0.3
Sambucus racemosa L.

Red-berried Elder

4

RUBIACEAE
Galium palustre agg.

Marsh Bedstraw

2

0.4

2

5 2 5
COMPOSITAE
Arctium minus Bernh.

Lesser Burdock

2

+

0.6

+ +
Cirsium arvense (L.) Scop.

Field Thistle

3

3

5

1 0.4 +
Cirsium palustre (L.) Scop.

Marsh Thistle

9

5

7

11 10 6
Cirsium vulgare (Savi) Ten.

Spear Thistle

+

Eupatorium cannabinum L.

Hemp-agrimony

1

2

1

3 2 2
Hypochoeris radicata L.

Cat’s Ear

0.2

5

5

6 10 10
Leontodon autumnalis L.

Autumn Hawkbit

0.4 2
Pulicularia dysenterica (L.) Bernh.

Common Fleabane

+

0.9
Senecio jacobaea L.

Ragwort

+ + + + 0.9
Taraxacum officinale agg.

Dandelion

0.8 1 0.4 0.7 0.4 2
Number of Species *62 84 80 79 76 92

 

* The species numbers for 1996 are underestimated because some plants, particularly grasses and sedges were identified only to generic level and not species level.

 

Summary ... the data at a glance, with notes.

  • !20 different plant species have been recorded on the Heathland Restoration Project site over the 6 years from 1996 -2001.

    Heathlands are typically species-poor and are generally dominated by a few plant species specially adapted to survive the nutrient-poor, often dry, sandy soils characteristic of heathland habitats. The relatively large number of species recorded on the project site reflects the fact that it is a habitat in flux, where many colonizing species are competing with each other. As time passes, the number of plant species is likely to decrease, as the heath species most suited to the microconditions of the site begin to dominate and out-compete the more opportunistic species.

  • Approximately 15% of the recorded species occurred with a percentage frequency of 20% or more over the whole project site. This means that the majority of the species recorded above are neither abundant, nor widespread on the site. However, some species are quite abundant in some sections of the site and not in others. This means that when their percentage frequency of occurrence is averaged out over the whole site, they appear less common than they actually are in certain areas of the project site. This includes species such as Tormentil and Bell Heather.

  • Heather and Bell Heather have been part of the naturally regenerating plant community from the beginning. Heather has gradually increased on the site, reaching a relatively steady percentage frequency of occurrence over the past 3 years, as has Bell Heather. This stands at approximately 20% frequency for Heather and approximately 6% frequency for Bell Heather.

  • Other characteristic heath species such as Tormentil, Heath Speedwell and the moss, Hypnum jutlandicum are slowly increasing in abundance. Typical heath grasses such as Bristle Bent have also begun to colonize the site.

  • European Gorse, which is a characteristic colonizer of areas of disturbed ground, is well established on the project site. Western Gorse, which is typical of heaths in this geographical region (Devon, SW England) has begun to appear on the site.

  • Silver Birch, which is the pioneer tree species most involved in the natural succession of heathland to woodland, has occurred with an approximate frequency of 30 - 40% across the whole site over the past 5 years.

  • It can clearly be seen that pioneer species such as Foxglove were initially quite abundant but have declined with time, as would be expected.

 

 

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